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Something dreadful has happened; the Flower Ponies' sister, Hyacinth, is coming to stay. Also, the entire town is being overrun by mind-swapping robot crabs, which may at least be enough to make Hyacinth quit complaining about the dirt.
Subject #040496 - once a human, now a pegasus - searches for a way home. Meanwhile, the Star Bright Network has ideas of their own for her, and the Doctor finds himself caught in a battle to maintain reality.
A dalek appears in Equestria - not just any dalek, but the one that had been corrupted by Rose Tyler's DNA. It has a choice of life as a pony, or dying, and finds that it cannot bear to die a second time. Thus begins a strange adventure...
Death sends the Tenth Doctor somewhere he never could have expected. With new friends to make and ancient foes from two universes to fight, only one thing is certain: there's an awful lot of running left to do.