Applejack has always had a secret dream, one that some ponies would find strange for the tomboyish farmer. She's always wanted somepony to treat her to a special romantic night, while being treated like a real lady. Turns out all she had to do was have a certain rainbow-maned pegasus catch her in a dress. Such an awkward moment will lead them to a night neither will ever forget.

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Twilight and her five friends are transported during a botched spell, across dimensions to... Ponyville, Equestria? But not the same one that they knew. Sitting on a bench is Harpsy and Babar. The Doctoress avoids apples from an aspiring Applebuck. And in a library across town, six stallions join their forces as they try to decipher the appearance of filly look-alikes of themselves around Ponyville!

What would you do, if you met a pony who was at once your brother, and yet not? Applebuck wishes to reunite with his 'Cousin JayJay', and braves the dimensional divide to reach his goal. But being a small foal, how will he deal with a world in which everything is flipped? His friends, his family, and even... himself?